The dog ( Canis familiaris or Canis lupus familiaris) is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by . The dog was the first species to be domesticated by , over 14,000 years ago and before the development of agriculture. Due to their long association with humans, dogs have gained the ability to thrive on a starch-rich diet that would be inadequate for other Canidae.
Dogs have been bred for desired behaviors, sensory capabilities, and physical attributes. vary widely in shape, size, and color. They have the same number of bones (with the exception of the tail), powerful jaws that house around 42 teeth, and well-developed senses of smell, hearing, and sight. Compared to humans, dogs possess a superior sense of smell and hearing, but inferior visual acuity. Dogs perform many roles for humans, such as Hunting dog, Herding dog, Sled dog, Guard dog, Pet, Therapy dog, Service dog, and Police dog and the military.
Communication in dogs includes eye gaze, facial expression, vocalization, body posture (including movements of bodies and limbs), and gustatory communication (scents, pheromones, and taste). They mark their territories by urinating on them, which is more likely when entering a new environment. Over the millennia, dogs have uniquely adapted to human behavior; this adaptation includes being able to understand and communicate with humans. As such, the human–canine bond has been a topic of frequent study, and dogs' influence on human society has given them the sobriquet of "man's best friend".
The global dog population is estimated at 700 million to 1 billion, distributed around the world. The dog is the most popular pet in the United States, present in 34–40% of households. Developed countries make up approximately 20% of the global dog population, while around 75% of dogs are estimated to be from developing countries, mainly in the form of feral and community dogs.
Dogs, wolves, and have sometimes been classified as separate species. In 1758, the Swedish botanist and zoologist Carl Linnaeus assigned the genus name Canis (which is the Latin word for "dog") to the domestic dog, the wolf, and the golden jackal in his book, Systema Naturae. He classified the domestic dog as Canis familiaris and, on the next page, classified the grey wolf as Canis lupus. Linnaeus considered the dog to be a separate species from the wolf because of its upturning tail ( cauda recurvata in Latin term), which is not found in any other canid. In the 2005 edition of Mammal Species of the World, mammalogist W. Christopher Wozencraft listed the wolf as a wild subspecies of Canis lupus and proposed two additional subspecies: familiaris, as named by Linnaeus in 1758, and dingo, named by Meyer in 1793. Wozencraft included hallstromi (the New Guinea singing dog) as another name (junior synonym) for the dingo. This classification was informed by a 1999 mitochondrial DNA study.
The classification of dingoes is disputed and a political issue in Australia. Classifying dingoes as wild dogs simplifies reducing or controlling dingo populations that threaten livestock. Treating dingoes as a separate species allows conservation programs to protect the dingo population. Dingo classification affects wildlife management policies, legislation, and societal attitudes. In 2019, a workshop hosted by the IUCN/Species Survival Commission's Canid Specialist Group considered the dingo and the New Guinea singing dog to be feral dog Canis familiaris. Therefore, it did not assess them for the IUCN Red List of threatened species.
DNA sequences show that all ancient and modern dogs share a common ancestry and descended from an ancient, extinct wolf population that was distinct from any Wolf lineage. Some studies have posited that all living wolves are more closely related to each other than to dogs, while others have suggested that dogs are more closely related to modern Eurasian wolves than to American wolves.
The dog is a domestic animal that likely travelled a Commensalism pathway into domestication (i.e. humans initially neither benefitted nor were harmed by wild dogs eating refuse from their camps). The questions of when and where dogs were first domesticated remains uncertain. Genetic studies suggest a domestication process commencing over 25,000 years ago, in one or several wolf populations in either Europe, the high Arctic, or eastern Asia. In 2021, a literature review of the current evidence Inference that the dog was domesticated in Siberia 23,000 years ago by ancient North Siberians, then later dispersed eastward into the Americas and westward across Eurasia, with dogs likely accompanying the first humans to inhabit the Americas. Some studies have suggested that the extinct Japanese wolf is closely related to the ancestor of domestic dogs.
In 2018, a study identified 429 genes that differed between modern dogs and modern wolves. As the differences in these genes could also be found in ancient dog fossils, these were regarded as being the result of the initial domestication and not from recent breed formation. These genes are linked to neural crest and central nervous system development. These genes affect embryogenesis and can confer tameness, smaller jaws, floppy ears, and diminished craniofacial development, which distinguish domesticated dogs from wolves and are considered to reflect domestication syndrome. The study concluded that during early dog domestication, the initial selection was for behavior. This trait is influenced by those genes which act in the neural crest, which led to the phenotypes observed in modern dogs.
Compared to the dog's wolf-like ancestors, selective breeding since domestication has seen the dog's skeleton increase in size for larger types such as and miniaturised for smaller types such as ; dwarfism has been selectively bred for some types where short legs are preferred, such as and . Most dogs naturally have 26 vertebrae in their tails, but some with Natural bobtail have as few as three.
The dog's skull has identical components regardless of breed type, but there is significant Cephalic index between types. The three basic skull shapes are the elongated dolichocephalic type as seen in , the intermediate mesocephalic or mesaticephalic type, and the very short and broad brachycephalic type exemplified by mastiff type skulls. The jaw contains around 42 teeth, and it has evolved for the consumption of flesh. Dogs use their carnassial teeth to cut food into bite-sized chunks, more especially meat.
Dogs' vision is Dichromacy; their visual world consists of yellows, blues, and grays. They have difficulty differentiating between red and green, and much like other mammals, the dog's eye is composed of two types of cone cells compared to the human's three. The divergence of the eye axis of dogs ranges from 12 to 25°, depending on the breed, which can have different retina configurations. The fovea centralis area of the eye is attached to a Axon, and is the most sensitive to . Additionally, a study found that dogs' visual acuity was up to eight times less effective than a human, and their ability to discriminate levels of brightness was about two times worse than a human.
While the human brain is dominated by a large visual cortex, the dog brain is dominated by a large olfactory system. Dogs have roughly forty times more smell-sensitive receptors than humans, ranging from about 125million to nearly 300million in some dog breeds, such as . This sense of smell is the most prominent sense of the species; it detects chemical changes in the environment, allowing dogs to pinpoint the location of mating partners, potential stressors, resources, etc. Dogs also have an acute sense of hearing up to four times greater than that of humans. They can pick up the slightest sounds from about compared to for humans.
Dogs have stiff, deeply embedded hairs known as that sense atmospheric changes, vibrations, and objects not visible in low light conditions. The lower most part of whiskers hold more receptor cells than other hair types, which help in alerting dogs of objects that could collide with the nose, ears, and jaw. Whiskers likely also facilitate the movement of food towards the mouth.
A study suggests that dogs show asymmetric tail-wagging responses to different emotive stimuli. "Stimuli that could be expected to elicit approach tendencies seem to be associated with a higher amplitude of tail-wagging movements to the right side". Dogs can injure themselves by wagging their tails forcefully; this condition is called kennel tail, happy tail, bleeding tail, or splitting tail. In some , the tail is traditionally docked to avoid injuries. Some dogs can be born without tails because of a DNA variant in the T gene, which can also result in a congenitally short (bobtail) tail. Tail docking is opposed by many veterinary and animal welfare organisations such as the American Veterinary Medical Association and the British Veterinary Association. Evidence from veterinary practices and showed that around 500 dogs would need to have their tail docked to prevent one injury.
Common dog parasites are lice, , fly larvae, , , , , and coccidia. Taenia is a notable genus with 5 species in which dogs are the definitive host. Additionally, dogs are a source of zoonoses for humans. They are responsible for 99% of rabies cases worldwide; however, in some developed countries such as the UK, rabies is absent from dogs and is instead only transmitted by bats. Other common zoonoses are hydatid disease, leptospirosis, pasteurellosis, ringworm, and toxocariasis. Common infections in dogs include canine adenovirus, canine distemper virus, canine parvovirus, leptospirosis, canine influenza, and canine coronavirus. All of these conditions have vaccines available.
Dogs are the companion animal most frequently reported for exposure to . Most poisonings are accidental and over 80% of reports of exposure to the ASPCA animal poisoning hotline are due to oral exposure. The most common substances people report exposure to are: pharmaceuticals, toxic foods, and rodenticides.
Neoplasia is the most common cause of death for dogs. Other common causes of death are heart failure and renal failure. Their pathology is similar to that of humans, as is their Immune response to treatment and their outcomes. Genes found in humans to be responsible for disorders are investigated in dogs as being the cause and vice versa.
In a 2024 UK study analyzing 584,734 dogs, it was concluded that purebred dogs lived longer than crossbred dogs, challenging the previous notion of the latter having the higher life expectancies. The authors noted that their study included "" as crossbred and that purebred dogs were typically given better care than their crossbred counterparts, which likely influenced the outcome of the study. Other studies also show that fully mongrel dogs live about a year longer on average than dogs with pedigrees. Furthermore, small dogs with longer muzzles have been shown to have higher lifespans than larger medium-sized dogs with much more depressed muzzles. For free-ranging dogs, less than 1 in 5 reach sexual maturity, and the median life expectancy for feral dogs is less than half of dogs living with humans.
Neutering is the most common surgical procedure in dogs less than a year old in the US and is seen as a control method for overpopulation. Neutering often occurs as early as 6–14 weeks in shelters in the US.
Neutering is less common in most European countries, especially in Nordic countries—except for the UK, where it is common. In Norway, neutering is illegal unless for the benefit of the animal's health (e.g., ovariohysterectomy in case of ovarian or uterine neoplasia). Some European countries have similar laws to Norway, but their wording either explicitly allows for neutering for controlling reproduction or it is allowed in practice or by contradiction through other laws. Italy and Portugal have passed recent laws that promote it. Germany forbids early age neutering, but neutering is still allowed at the usual age. In Romania, neutering is mandatory except for when a pedigree to select breeds can be shown.
Most domestic animals were initially bred for the production of goods. Dogs, on the other hand, were selectively bred for desirable behavioral traits.
Dogs have a natural instinct called prey drive (the term is chiefly used to describe Dog training' habits) which can be influenced by breeding. These instincts can drive dogs to consider objects or other animals to be prey or drive possessive behavior. These traits have been enhanced in some breeds so that they may be used to hunt and kill vermin or other pests. Puppies or dogs sometimes bury food underground. One study found that wolves outperformed dogs in finding food caches, likely due to a "difference in motivation" between wolves and dogs. Some puppies and dogs engage in Coprophagia out of habit, stress, for attention, or boredom; most of them will not do it later in life. A study hypothesizes that the behavior was inherited from wolves, a behavior likely evolved to lessen the presence of intestinal parasites in dens. Most dogs can swim. In a study of 412 dogs, around 36.5% of the dogs could not swim; the other 63.5% were able to swim without a trainer in a swimming pool. A study of 55 dogs found a correlation between swimming and 'improvement' of the hip Osteoarthritis.
One study of canine cognitive abilities found that dogs' capabilities are similar to those of , chimpanzees, or . One study of 18 household dogs found that the dogs could not distinguish food bowls at specific locations without distinguishing cues; the study stated that this indicates a lack of spatial memory. A study stated that dogs have a visual sense for number. The dogs showed a ratio-dependent activation both for numerical values from 1–3 to larger than four.
Dogs demonstrate a theory of mind by engaging in deception. Another experimental study showed evidence that Australian can outperform domestic dogs in non-social problem-solving, indicating that domestic dogs may have lost much of their original problem-solving abilities once they joined humans. Another study showed that dogs stared at humans after failing to complete an impossible version of the same task they had been trained to solve. Wolves, under the same situation, avoided staring at humans altogether.
Humans communicate with dogs by using vocalization, hand signals, and body posture. With their acute sense of hearing, dogs rely on the auditory aspect of communication for understanding and responding to various cues, including the distinctive barking patterns that convey different messages. A study using functional magnetic resonance imaging (fMRI) has shown that dogs respond to both vocal and nonvocal voices using the brain's region towards the temporal pole, similar to that of humans' brains. Most dogs also looked significantly longer at the face whose expression matched the valence of vocalization. A study of Caudate nucleus shows that dogs tend to respond more positively to social rewards than to food rewards.
and have also been known to attack dogs. In particular, are known to have a preference for dogs and have been recorded to kill and consume them, no matter their size.
Of the twenty-one amino acids common to all life forms (including selenocysteine), dogs cannot synthesize ten: arginine, histidine, isoleucine, leucine, lysine, methionine, phenylalanine, threonine, tryptophan, and valine. Like cats, dogs require arginine to maintain nitrogen balance. These nutritional requirements place dogs halfway between carnivores and omnivores.
Dogs were introduced to Antarctica as . Starting practice in December 1993, dogs were later outlawed by the Protocol on Environmental Protection to the Antarctic Treaty international agreement due to the possible risk of spreading infections.
Products such as dog-training books, classes, and television programs target dog owners. Some dog-trainers have promoted a dominance model of dog-human relationships. However, the idea of the "alpha dog" trying to be dominant is based on a controversial theory about wolf packs. It has been disputed that "trying to achieve status" is characteristic of dog-human interactions. Human family members have increased participation in activities in which the dog is an integral partner, such as dog dancing and dog yoga.
According to statistics published by the American Pet Products Manufacturers Association in the National Pet Owner Survey in 2009–2010, an estimated 77.5 million people in the United States have pet dogs. The source shows that nearly 40% of American households own at least one dog, of which 67% own just one dog, 25% own two dogs, and nearly 9% own more than two dogs. The data also shows an equal number of male and female pet dogs; less than one-fifth of the owned dogs come from Animal shelter.
Eating dog meat is a Taboo in most parts of the world, though some still consume it in Modern era. It is still consumed in some East Asian countries, including China, Vietnam, Korea, Indonesia, and the Philippines. An estimated 30 million dogs are killed and consumed in Asia every year. China is the world's largest consumer of dogs, with an estimated 10 to 20 million dogs killed every year for human consumption. In Vietnam, about 5 million dogs are slaughtered annually. In 2024, China, Singapore, and Thailand placed a ban on the consumption of dogs within their borders. In some parts of Poland and Central Asia, dog fat is reportedly believed to be beneficial for the lungs. Proponents of eating dog meat have argued that placing a distinction between livestock and dogs is Western hypocrisy and that there is no difference in eating different animals' meat.
There is a long history of dog meat consumption in South Korea, but the practice has fallen out of favor. A 2017 survey found that under 40% of participants supported a ban on the distribution and consumption of dog meat. This increased to over 50% in 2020, suggesting changing attitudes, particularly among younger individuals. In 2018, the South Korean government passed a bill banning restaurants that sell dog meat from doing so during that year's Winter Olympics. On 9 January 2024, the South Korean parliament passed a law banning the distribution and sale of dog meat. It will take effect in 2027, with plans to assist dog farmers in transitioning to other products. The primary type of dog raised for meat in South Korea has been the Nureongi. In North Korea where meat is scarce, eating dog is a common and accepted practice, officially promoted by the government.
Toxocara canis (dog roundworm) eggs in dog feces can cause toxocariasis. It is estimated that nearly 14% of people in the United States are infected with Toxocara; about 10,000 cases are reported each year. Untreated toxocariasis can cause retinal damage and decreased vision. Dog feces can also contain that cause cutaneous larva migrans in humans.
A 2005 paper by the British Medical Journal states:
One study indicated that wheelchair-users experience more positive social interactions with strangers when accompanied by a dog than when they are not. In a 2015 study, it was found that having a pet made people more inclined to foster positive relationships with their neighbors. In one study, new guardians reported a significant reduction in minor health problems during the first month following pet acquisition, which was sustained through the 10-month study.
Using dogs and other animals as a part of therapy dates back to the late-18th century, when animals were introduced into mental institutions to help socialize patients with . Animal-assisted intervention research has shown that animal-assisted therapy with a dog can increase smiling and laughing among people with Alzheimer's disease.
In mythology, dogs often appear as pets or as watchdogs. Stories of dogs guarding the gates of the underworld recur throughout Indo-European mythologies and may originate from Proto-Indo-European traditions. In Greek mythology, Cerberus is a three-headed, dragon-tailed watchdog who guards the gates of Hades. Dogs also feature in association with the Greek goddess Hecate. In Norse mythology, a dog called Garmr guards Hel, a realm of the dead. In Persian mythology, two four-eyed dogs guard the Chinvat Bridge. In Welsh mythology, Cŵn Annwn guards Annwn. In Hindu mythology, Yama, the god of death, owns two watchdogs named Shyama and Sharvara, which each have four eyes—they are said to watch over the gates of Naraka. A black dog is considered to be the vahana (vehicle) of Bhairava (an incarnation of Shiva).
In Christianity, dogs represent faithfulness. Within the Roman Catholic denomination specifically, the iconography of Saint Dominic includes a dog after the saint's mother dreamt of a dog springing from her womb and became pregnant shortly after that. As such, the Dominican Order (Ecclesiastical Latin: Domini canis) means "dog of the Lord" or "hound of the Lord". In Christian folklore, a church grim often takes the form of a black dog to guard Christian churches and their from sacrilege. Jewish law does not prohibit keeping dogs and other pets but requires Jews to feed dogs (and other animals that they own) before themselves and to make arrangements for feeding them before obtaining them. The view on dogs in Islam is mixed, with some schools of thought viewing them as unclean, although Khaled Abou El Fadl states that this view is based on "pre-Islamic Arab mythology" and "a tradition ... falsely attributed to the Prophet". The Sunni Maliki school jurists disagree with the idea that dogs are unclean.
Breeds
There are around 450 official [[dog breeds]], the most of any mammal. Dogs began diversifying in the [[Victorian era]], when humans took control of their natural selection. Most breeds were derived from small numbers of founders within the last 200 years. Since then, dogs have undergone rapid phenotypic change and have been subjected to artificial selection by humans. The skull, body, and limb proportions between breeds display more phenotypic diversity than can be found within the entire order of [[carnivore]]s. These breeds possess distinct traits related to morphology, which include body size, skull shape, tail phenotype, fur type, and colour. As such, humans have long used dogs for their desirable traits to complete or fulfill a certain work or role. Their behavioural traits include guarding, herding, hunting, retrieving, and scent detection. Their personality traits include hypersocial behavior, boldness, and aggression. Present-day dogs are dispersed around the world. An example of this dispersal is the numerous modern breeds of European lineage during the Victorian era.
File:Dog morphological variation.png|Morphological variation in six dogs
File:Dog coat variation.png|Phenotype in four dogs
Anatomy and physiology
Size and skeleton
Senses
Coat
Dewclaw
Tail
Health
Lifespan
Reproduction
Neutering
Inbreeding depression
Behavior
Nursing
Intelligence
Communication
Ecology
Population
Competitors
Diet
Despite being an omnivore, dogs are only able to conjugate bile acid with taurine. They must get vitamin D from their diet.
Range
Roles with humans
Pets
Workers
Shows and sports
Dogs as food
Health risks
Health benefits
Recent research has failed to support earlier findings that pet ownership is associated with a reduced risk of cardiovascular disease, a reduced use of general practitioner services, or any psychological or physical benefits on health for community dwelling older people. Research has, however, pointed to significantly less absenteeism from school through sickness among children who live with pets.
Health benefits of dogs can result from contact with dogs in general, not solely from having dogs as pets. For example, when in a pet dog's presence, people show reductions in cardiovascular, behavioral, and psychological indicators of anxiety and are exposed to Probiotic, which can protect against allergies and autoimmune diseases (according to the hygiene hypothesis). Other benefits include dogs as social support.
Cultural importance
Terminology
See also
Bibliography
External links
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